The aerodynamics of revolving wings I. Model hawkmoth wings
James R. Usherwood* and
Charles P. Ellington
Department of Zoology, University of Cambridge, Downing Street,
Cambridge CB2 3EJ, UK
* Present address: Concord Field Station, MCZ, Harvard University, Old Causeway
Road, Bedford, MA 01730, USA

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Fig. 1 . Propeller body (A) and plan and side views (B) of the complete `see-saw'
propeller rig. Roman numerals identify parts of the propeller body. i, sting
for attachment to wings; ii, propeller head; iii, smoke chamber (smoke in this
chamber feeds into the hollow shaft and up to the propeller head); iv,
cut-away section showing torque strain gauges (electrical connections run down
the hollow axle); v, strain gauge bridge supply and first-stage amplification
by electronics rotating with axle; vi, electrical contacts on multi-wiper
slip-rings carrying power and strain gauge signal; vii, gearbox; viii, motor;
ix, tachometer.
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Fig. 2. Standard (A) and `sawtooth' (B) hawkmoth planforms.
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Fig. 3 . Typical voltage signals for a single run at high angle of attack
(A) and response of the vertical force transducer to the addition, and then
removal, of 5, 10 and 20 g before (B) and after (C) filtering. In A, the top
(green) line shows the tachometer trace, the middle (blue) line the torque
signal and the bottom (red) line the vertical force signal. The wings were
started after 10s. Vertical dotted lines identify five oscillations due to the
lightly damped `mass-spring' system (i) inherent in the vertical transducer
design, and one cycle due to a mass imbalance of the wings (ii) during a
complete revolution.
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Fig. 4 . Averaged horizontal Ch (A) and vertical
Cv (B) force coefficients plotted against angle of
revolution for standard hawkmoth wings over the `abbreviated' range of angle
of attack . Underlying grey panels show the averaging period for
`early' (narrower panel) and `steady' (broader) pools.
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Fig. 6. Horizontal Ch (A) and vertical Cv
(B) force coefficients and the polar diagram (C) for standard hawkmoth wings
under `early' and `steady' conditions. Error bars in A and B show ±1
S.E.M., N=4-10. ', angle of incidence.
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Fig. 7. Horizontal Ch (A) and vertical Cv
(B) force coefficients and the polar diagram (C) for the `leading-edge' range
under `early' and `steady' conditions. Underlying grey lines show `early'
(higher) and `steady' (lower) values for standard hawkmoth wings and represent
0° twist and 0% camber. ', angle of incidence.
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Fig. 8. Horizontal Ch (A) and vertical Cv
(B) force coefficients and the polar diagram (C) for hawkmoth wings with a
range of twist under `early' and `steady' conditions. Underlying grey lines
show `early' (higher) and `steady' (lower) values for standard hawkmoth wings.
', angle of incidence.
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Fig. 9. Horizontal Ch (A) and vertical Cv
(B) force coefficients and the polar diagram (C) for hawkmoth wings with a
range of camber under `early' and `steady' conditions. Underlying grey lines
show `early' (higher) and `steady' (lower) values for standard hawkmoth wings.
', angle of incidence.
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Fig. 10. Polar diagram showing results from three models for determining the profile
drag coefficient CD,pro and the lift coefficient
CL from the `steady' data represented by the lower yellow
line. A good model would result in values close to, or slightly above, those
of the `early' conditions represented by the upper yellow line. Data are
`pooled' values for all wings in the `leading-edge' series.
Ch, horizontal force coefficient; Cv,
vertical force coefficient; , geometric angle of attack.
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© The Company of Biologists Ltd 2002