First published online December 14, 2006
Journal of Experimental Biology 210, 56-64 (2007)
Published by The Company of Biologists 2007
doi: 10.1242/jeb.02618
Echolocation signals of wild harbour porpoises, Phocoena phocoena
Anne Villadsgaard1,*,
Magnus Wahlberg1,2 and
Jakob Tougaard3
1 Department of Zoophysiology, Institute of Biological Sciences, University
of Aarhus, C. F. Møllers Alle, DK-8000 Aarhus C., Denmark
2 Fjord & Bælt, Margrethes Plads 4, DK-5300 Kerteminde,
Denmark
3 National Environmental Research Institute, Fredriksborgvej 399, DK-4000
Roskilde, Denmark

View larger version (6K):
[in this window]
[in a new window]
|
Fig. 1. The experimental set up consists of a linear, vertical array of three
(A,B,D) or four (A-D) hydrophones connected via a conditioning box to
a digital recorder.
|
|

View larger version (8K):
[in this window]
[in a new window]
|
Fig. 2. (A) Harbour porpoise click with signal envelope (dotted line) and the -10
dB duration of the click (horizontal line). (B) Accumulated energy content (%)
in the click over time. The interval between the dotted lines is covering 95%
of the energy content of the signal.
|
|

View larger version (8K):
[in this window]
[in a new window]
|
Fig. 3. Results from the transmission loss measurements and acoustic ranging
calibration, using an omni-directional calibrated sound source emitting
harbour porpoise clicks at a known range from 3 m depth. (A) Received level
(dB re 1 µPa pp) as a function of measured range for the 4-hydrophone
array. The line is a transmission loss model consisting of spherical spreading
and absorption. (B) RMS error (see text) in acoustic localization as a
function of the measured range for the 4-hydrophone array and the 3-hydrophone
array. (C) The maximum variation in the transmission loss due to ranging
variation as a function of measured range for the 4-hydrophone array and the
3-hydrophone array.
|
|

View larger version (6K):
[in this window]
[in a new window]
|
Fig. 4. The estimated apparent source level of harbour porpoise clicks as a
function of the back-calculated range between the porpoise and the array.
|
|

View larger version (5K):
[in this window]
[in a new window]
|
Fig. 5. Power spectrum of a harbour porpoise click (FFT size 64, spectrum
interpolated with a factor 10, sampling rate 400 kHz, rectangular window).
|
|

View larger version (7K):
[in this window]
[in a new window]
|
Fig. 6. Histogram of inter-click intervals (ICIs) from wild harbour porpoises
(N=822; median 58 ms; s.d. 34 ms; bin width 10 ms).
|
|
© The Company of Biologists Ltd 2007