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Fig. 3. Schematics showing measurement techniques for determining physiological
cross-sectional area and theoretical maximum force production in the clavatus
muscle (CM) and deep protractor muscle (DPM). (A) Fiber angle measurements did
not apply to the DPM, as DPM fibers coursed parallel with the line of action
(broken blue line). (B) Lower inset shows that the two heads of the CM
separate once it is removed from the feeding apparatus. Upper inset
demonstrates how pennation angles were measured. Fiber angles
are
identical near the muscle's midline (line of action) and begin to vary
slightly as they course posteriorly, therefore fiber angles were measured near
the muscle's midline. Muscle length L was used as a proxy for
determining fiber length because mean fiber lengths in the DPM and CM were
approximately equal to respective muscle lengths. Darkened areas on the
feeding apparatuses indicate positions of the CM and DPM in dorsal and ventral
views, respectively. Individual CM and DPM (from insets) are not drawn to
scale.